To prepare for a hiking trip in the Appalachian mountains you buy several field guides for the region from a used bookstore. You notice that the older guides list a single species of tiger swallowtail butterfly Papilio glaucus and an Appalachian morph of this species. However the newest guide list two separate species P. glaucus and P. appalachiensis. The authors of this guide state that the biological species concept was used in determining their classifications. What do you think led them to list two separate species of tiger swallowtail?